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Family Colletidae - Cellophane, Plasterer, Masked, and Allied Bees

Colletes inaequalis  - Colletes inaequalis Unidentified Insect 02_2012 - Colletes Colletes? - Colletes ochraceus Colletidae, Beauty Bee, lateral - Colletes - female Cellophane Bee - Colletes is this an Eastern Masked Bee (Hylaeus affinis)? - Hylaeus modestus - male Hylaeus confluens - Hylaeus - female Xylocopa ♂ - Ptiloglossa arizonensis
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon (Aculeata - Ants, Bees and Stinging Wasps)
No Taxon (Apoidea (clade Anthophila) - Bees)
Family Colletidae (Cellophane, Plasterer, Masked, and Allied Bees)
Other Common Names
Masked Bees
Numbers
ca. 160 spp. in 6 genera of 4 subfamilies in our area, >2,500 spp. in ~90 genera of 8 subfamilies worldwide(1)
Subfamilies:
Colletinae: ca. 100 spp. in a single genus in our area, almost 500 spp. in 2 genera total (the second genus is in S.America south of the equator)(1)
Diphaglossinae: 7 spp. in 3 genera in our area (all in the Caupolicanini), 130 spp. in 11 genera of 3 tribes total; this is a New World group, almost exclusively neotropical(1)
Hylaeinae: in our area, ~50 spp. in a single genus, 934 spp. in 7 genera worldwide; about ¾ of the global diversity is in the Old World; all the genera except the cosmopolitan Hylaeus are restricted to Australasia (~350 spp. are Australasian, too)(1)
Paracolletinae: a single species in our area, 430 spp. in 46 genera total; ~2/3 of the spp. are Australasian, the rest, Neotropical(1)
Overview of our fauna (* –taxa not yet in the guide)
Family Colletidae
Subfamily Colletinae Colletes
Subfamily Diphaglossinae
Identification
Jugal lobe in HW longer than submedian cell. Basal vein straight or nearly so. One subantennal suture below each antennal socket. Glossa short, bilobed or truncate. 2nd and 3rd submarginal cells are roughly the same size (in Andrenidae, 2nd one is distinctly smaller)(2)(3)
Guide to eastern genera in (4)
OVERVIEW OF GENERA
Subfamily Colletinae
Colletes - Cellophane bees
Caupolicana - Fork-tongued Bees
Ptiloglossa - Feather-tongued Bees
Zikanapis - Zikan's Feather-tongued Bees
Subfamily Hylaeinae
Hylaeus - Masked Bees
Range
Worldwide, most speciose in S.America and Australia
Life Cycle
Plasterer bees are the first to appear in spring.(5)
Remarks
Euryglossinae and Hylaeinae lack external pollen-carrying apparatus (scopa), they carry pollen in their crops. Plasterer bees use a secretion to plaster the walls of their nests. This dries into a cellophane-like lining. The lining envelopes the store of pollen and nectar, keeping it fresh in the damp, underground chambers.(5)